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Liberia Cook , Nigeria Cook , Ghana this study. Piona seyrigi continentalis Lundblad, New records. One of the males from the fish pond along road Elubo-Axim has around 35 pairs of acetabula, all other males have less than 30 pairs of acetabula. According to Cook the only reliable character to separate the nominate form and continentalis is the number of acetabula, in the seyrigi seyrigi and in seyrigi continentalis.
This study shows there can be an overlap in the number of acetabula, and that this character is not always reliable. Uganda Lundblad , Liberia Cook , Ghana this study. The assignment of this species is uncertain. Cook tentatively assigned specimens from Liberia to this species, which was described by Walter based on the female only.
The specimens from Ghana match the description of Cook well. Angola Walter , Liberia Cook , Ghana this study. Encentridophorus Encentridophorus vietsi Bader, New records. Viets assigned specimens from Cameroon to E. However, Encentridophorus females show few species-specific characters, and therefore I agree with Bader that the assignment of Viets is questionable.
Bader described his specimens from Cameroon as E. In my opinion the male from the Gambia and the specimens from Viets belong all to E. Bader was the opinion that the specimens reported by Walter from Chad do not belong to E. Bader based this opinion on the measurements of leg segments and the number of swimming setae. Subfamily Pionatacinae K. Viets Neumania Alloneumania marginata K.
Viets, — nov. Ecpolus dorsofenestratus Lundblad, — new syn. Cook suggested that Neumania marginata K. Viets, might belong to the subgenus Alloneumania. Males of Neumania dorsofenestrata Lundblad and females of N. Therefore, the suggestion of Cook was correct. I do not see any differences between the males from Ghana and Congo, and therefore I propose to synonymize N. Young males do not have a dorsal shield. Walter reported this species from Niger and Burkina Faso.
His illustration of the female shows a narrow strip of secondary sclerotization along Cx-IV. However, N. In my opinion the specimens of Walter must be assigned to N. The female of this species has a genital field which is very similar to that of N. Therefore, it is not clear to which species their material belongs. Cameroon Viets , the Gambia, Ghana this study.
Neumania Allolemienia falcifera Cook, female. A — venter; B — palp; C — IV-leg Female: Integument smooth, idiosoma dorsally long and wide, ventrally long. Apodemes of anterior coxae reaching to anterior part of Cx-IV. Venter without secondary sclerotization. Gonopore long. Genital plates rounded, long, somewhat longer than wide, each plate with acetabula.
Between Cxgl-4 and genital field four pairs of setae lying free in the integument. Length of P1—5: 42, , 72, , P4 with a large ventral setal tubercle, more medially two more smaller setal tubercles present Figure 17B. Length of I-leg-4—6: , , ; grooved setae very long. Claws of legs relatively large, e. IV-leg-5 with one trifurcate seta, other trifurcate setae lost. Swimming setae lost, but probably IV-leg-5 and IV-leg-4 with four swimming setae.
The male specimen from the Gambia differs somewhat from the holotype and only known specimen thus far. The large claw of II-leg-6 is long in the holotype , and IV-leg-3 and IV-leg-4 have only one trifurcate ventral seta versus IV-leg-3 with two or three and IV-leg-4 with three in the holotype. As nothing is known about the variation in these characters, I assign the specimen from West Africa for the time being to falcifera.
The female has not been described previously. Uganda Cook , the Gambia, Ghana this study. Neumania Soarella fusiformis n. A — dorsum; B — venter; C — palp. Apodemes of anterior coxae short, extending to Cx-III; post-genital sclerite large; excretory pore not on a tubercle.
Female: Integument with relatively large spiny setae. Idiosoma with large spindle-shaped tubercles. Dorsum with a pair of small platelets Figure 18A. Coxal field small, apodemes of anterior coxae short, extending to Cx-III. Genital field wide, genital plates 58 long with seven pairs of acetabula. Pre-genital sclerite much smaller the post-genital sclerite, the latter relatively large, wide Figure 18B. Excretory pore not on a tubercle.
Length of P 20, 69, 31, 56, P4 anteroventrally with a short, blunt seta Figure 18C. Length of I-leg-4—6: 60, 70, IV-leg-3—6 ventrally with 2, 2, 3, 1 pectinate setae, respectively. Named for the large spindle-shaped tubercles. The new species is somewhat similar to N. Neumania Soarella ghanaensis n. A — dorsum; B — venter; C — palp; D — venter. Male: Acetabula in two distinctly separated groups, anteriorly pairs, posteriorly pairs; one of the claws of P5 large.
Female: Acetabula as in male, but genital field with a lateral extension. Integument covered with long, spiny setae. Glandularia, postocularia and excretory pore on relatively short tubercles Figure 19A. Apodemes of anterior coxae extending to anterior part of Cx-IV. Posterior margin of Cx-IV with a short extension.
Cx-III anterolaterally with an extension. Male: Idiosoma long and wide, ventrally long. Gonopore 60 long and 34 wide, genital field wide. Acetabula in two groups, a group of 4or 5 anterior acetabula distinctly separated from posterior pairs Figure 19B.
Length of P1—5: 22, 92, 38, 82, P2 relatively long, P4 relatively stocky, the latter segment anteroventrally with a blunt seta, dorsally with two long, curved setae. P5 with one large claw Figure 19C. Grooved or fluted setae of first leg larger than those of second and third legs.
IV-leg-3, -4 and -5 ventrally with two, three and four pectinate setae, respectively. I-leg-5 and II-leg-5 with one and two short swimming setae, respectively. Dorsum with two small platelets, 16 in diameter. Genital field with two fused groups of acetabula, anteriorly pairs of acetabula well separated from the posterior acetabula, posteriorly pairs of acetabula; genital field with a lateral extension with three pairs of acetabula Figure 19D.
Length of P1—5: 22, 88, 42, 72, P2 slender as in male, but P5 with the large claw. Legs as in male, but IV-leg-5 with two or three swimming setae. Named after the country where the new species was collected. The new species has smaller tubercles, and three of the dorsal tubercles are located medially in ecphyma all tubercles are located near the lateral idiosoma margin. Moreover, the lateral extension of the genital field is absent in N.
Neumania Neumania liberiensis Cook, Figure 20 Figure Neumania Neumania liberiensis Cook, dorsum male. The male matches the description of Cook , apart from the apodemes of the anterior coxae. Cook was uncertain about the assignment of the female, but in my opinion this is correct.
Females have the venter with less secondary sclerotization compared to the male, and the apodemes of the anterior coxae extend to the middle of Cx-IV. Some measurements are given below. Males: Integument smooth. Females: Idiosoma dorsally long and wide, ventrally long. Neumania Leptopterotrichophorus manokensis K. Viets , Liberia Cook , Ghana this study. Neumania Neumania marginata K.
Cameroon Viets , South Africa K. Neumania Neumania paucipora Koenike, New records. Neumania Neumania reticulata K. Neumania Lemienia separata Cook, New records. Neumania Soarella thori Viets, New records. Genus Nyangalla K. Viets, Nyangalla Ecpolopella acuticaudata K. Figure 21 Figure Nyangalla Ecpolopella acuticaudata K. Viets, dorsum female. Female: Idiosoma brownish, dorsally long and wide, ventrally long.
Dorsal shield consisting of four pairs of platelets and an unpaired central platelet, flanked by four pairs of smaller lateral platelets and one pair of very small platelets. Glandularia are present on the four pairs of lateral platelets, the central platelet and the pair posterior to the central platelet.
The unpaired central platelet is long. Venter, palp and legs as in N. Male: Posterior margin of cauda with spiny setae. Previously this taxon was considered a subspecies of N. As already mentioned by Viets , the two taxa can be found on the same locality, and therefore, they cannot belong to the same species. Also, in Ghana the two taxa co-occur. Moreover, there are distinct differences between the two taxa. The female of N. The female of the N. The male of N.
Given the large differences between the two taxa, in males as well as in females, and the co-occurrence of the two taxa, I propose to consider N. Viets , Liberia Cook , the Gambia, Ghana this study. Nyangalla Ecpolopella peltophora K. Nyangalla Ecpolopella tesselata Daday, New records. The species has been described by Daday and Viets Therefore, I give only some measurements and additional characters. Male: Idiosoma long measured from anterior idiosoma margin till posterior margin of cauda , wide.
Posterior margin of cauda with spiny setae. Female: Idiosoma long and wide. Medial area between Cx-IV almost filled up completely by secondary sclerotization. Unionicola heversi n. A - venter; B — palp; C — palp; D — I-leg-4—6. Cx-II and Cx-III with anterolaterally with a pointed extension; palp stocky, especially P4, this segment with two setal tubercles and anteromedially a stout pointed seta, P5 with large claws, ventrally with two claws but only visible in a skew position.
Anterior coxae with short apodemes, extending to Cx-III. Posterior margin of Cx-IV with a short projection. Genital field with five pairs of acetabula on two pairs of platelets, all platelets with long posterior setae. Anterior platelets with two acetabula each, posterior pair with three acetabula each. Unfortunately, the genital field is distorted in the slide.
Palp stocky, especially P4, the latter segment with two setal tubercles and a short, stout pointed seta anteromedially. P5 with large claws, ventrally with two claws but only visible in a skew position as these are lying behind each other Figure 22B-C. Long setae of first leg not fluted or grooved Figure 22D.
IV-leg-5 with three, IV-leg-4 with two swimming setae. The palp with its large claws is somewhat similar to palps of U. Unionicola koenikei K. Outside Africa reported from Israel Por Unionicola latilaminata K. Unionicola megalopsis K. Viets , Ivory Coast Hevers , Ghana this study. Unionicola minuta K. Unionicola pollicigera K. Congo K. Some doubtful records published by Walter , , based on deutonymphs only. Arrenurus Arrenurus chappuisi Walter, New records.
Arrenurus chappuisi is close to A. I separated the two species based on the presence or absence of a two-lobed dorsal shield, and in lateral view the petiole is straight or upturned. With new material available of A. Sudan, Congo Bader , Ghana this study.
Arrenurus Megaluracarus chutteri ankasa n. A — dorsum; B — venter; C — palp; D - venter. Paratypes: one male, one female, same data as holotype RMNH. Cauda of male expanding laterally, posterior margin of cauda with a relatively wide median cleft which is widening posteriorly. Male: Idiosoma greenish, dorsally long and wide, ventrally long. Dorsal shield wide. Anterior idiosoma margin convex. Lateral idiosoma margin bulging anterior to the position where the genital plates extend onto the lateral part of the idiosoma.
Cauda expanding posterolaterally, in dorsal view lateral margins of cauda concave Figure 23A. Posterior margin of cauda with a relatively wide median cleft which is widening posteriorly; lateral to cleft a pair of small setae. Dgl-4 lying in an area with spiny setae. Gonopore 52 long. Genital plates long, extending onto lateral margins of idiosoma Figure 23B. Length of P1—5: 24, 56, 30, 60, P2 medially with three setae Figure 23C. IV-leg-4 with a short spur, IV-leg-6 slightly bowed.
Third and fourth legs with numerous swimming setae. Dorsal shield complete, long and wide. Anterior coxae extending beyond anterior idiosoma margin. Medial distance of Cx-IV smaller than width of one genital valve. Gonopore field wide, with anteriorly and posteriorly a small band of sclerotized patches; gonopore 92 long. Genital plates long and narrow, extending to lateral idiosoma margin Figure 23D. Length of P1—5: 27, 56, 26, 66, P2 medially with four setae.
Length of IV-lg-4—6: , , Named after Ankasa National Park, the name is a noun in apposition. The male of the new species differs from A. Viets, and A. The female of A. Arrenurus Megaluracarus chutteri longipes Smit, New records. Botswana Smit , Ghana this study. Arrenurus Micruracarus circulodorsalis n. Paratypes: four females one not sclerotized , same data as holotype RMNH. Other material. Male: Dorsum with a small, rounded dorsal shield, idiosoma posteriorly with a large hollow, posterior idiosoma margin with a closed cleft, gonopore not visible in ventral or posteromedial view.
Female: Genital plates extending halfway gonopore field and lateral idiosoma margin and widening laterally. Male: Idiosoma yellowish, dorsally long and wide, ventrally long. Anterior idiosoma margin straight.
Dorsal shield almost circular, long and wide, with the postocularia and Dgl Dgl-1 shifted posteriorly, somewhat posterior to level of Dgl-2 Figure 24A. No petiole present, although the structure illustrated at the posterior margin might suggest this, but this is just an interruption of the idiosoma pores. Idiosoma with a large hollow posteriorly, but this can be examined only in posteromedial view.
Gonopore not visible, not in ventral view, nor in posteromedial view. Posterior idiosoma margin straight, with a closed cleft, which widens anteriorly Figure 24B. Length of P1—5: 26, 60, 46, 72, P2 medially with three setae Figure 24C. Length of IV-leg , 98, I- and II-leg-6 with numerous small setae.
Legs with many swimming setae. Anterior idiosoma margin straight, posterolateral corners of idiosoma absent. Anterior coxae extending to or just beyond anterior idiosoma margin. Medial distance of Cx-IV longer than width of one genital valve. Gonopore field wide and without sclerotized patches, gonopore long. Genital field wide, genital plates widening laterally, extending halfway gonopore field and lateral idiosoma margin Figure 24D.
Excretory pore lying halfway gonopore field and posterior idiosoma margin. Length of P1—5: 32, 72, 58, 80, P3 medially with one seta. I- and II-leg-6 with less small setae compared to male. Named for the rounded dorsal shield of the male. The male is well characterized by its rounded dorsal shield, the absence of a petiole and the gonopore not visible in ventral or posteromedial view.
No other Afrotropical Arrenurus species shares this combination of characters. The female is well characterized by the shape of the genital plates. Arrenurus Micruracarus damkoehleri K. Arrenurus Megaluracarus geniculatus Koenike, female. Female: Idiosoma colour greenish. Anterior idiosoma margin slightly concave, posterolateral corners of idiosoma indistinct. Dorsal shield incomplete, wide. Medial distance of Cx-IV shorter than width of gonopore field.
Genital field wide, genital plates relatively broad, extending more or less halfway lateral idiosoma margin Figure 25A. Length of P1—5: 20, 60, 34, 68, P2 medially with three short setae; P4 stocky, anteroventrally rounded, anterior margin with a small extension Figure 25B. Legs with numerous swimming setae. The female of this species has not been described before. The small extension of the anterior margin of P4 is also present in the males of this study.
Madagascar, Ghana this study. Arrenurus Brevicaudaturus ghanaensis Cook, New records. Ghana Cook , this study. Arrenurus Brevicaudaturus neolaticodulus Cook, New records. Liberia Cook , Ghana Smit Arrenurus Brevicaudaturus oldhami Cook, New records.
Nigeria, Ghana Cook , this study. Arrenurus Rhinophoracarus praeacutus K. Cameroon, Ghana Cook , this study. Arrenurus Micruracarus tubifer Walter, New records. First records for Ghana. Genus Thoracophoracarus K. Viets, A genus with nine species known from Africa Gerecke Furthermore, one species known from Chile and one from Asia Smit Thoracophoracarus Thoracophoracarus kuehnei K.
With a straight posterior margin, a convex posterior margin and P2 with two setae, the specimen from Ghana matches the description given by Gerecke well. Thoracophoracarus Thoracophorurus petioluriger K. Viets, Figure 26A-C Figure Viets, female. Female: Idiosoma colour brownish. Anterior idiosoma margin strongly concave, excretory pore terminal lying in a concave posterior idiosoma margin.
Glandularia tubercles in posterior half of dorsum large Figure 26A. All suture lines of the coxae indistinct. Gonopore field 90 long, genital plates very indistinct and hardly discernible Figure 26B. Length of P1—5: 18, 42, 30, 50, P2 medially with three setae, P4 anteroventrally pointed Figure 26C. Length of IV-leg-4—6: 80, 84, As stated by Gerecke , attribution of males and females in arrenurids can be problematic due to the strong sexual dimorphism. This applies to the specimen of this study also.
Based on the strong concave anterior margin, assignment to T. However, due to the relative slender P4 and the large humps of the dorsum, I assigned this specimen to A. Females of the latter species, originally described as T. Viets, , are known from two specimens only collected in Cameroon. Moreover, there are some differences with the description given by Gerecke , i.
Therefore, an emended description is given of the female of petioluriger. Viets, A genus with numerous species known from the Afrotropical region. Djeboa expansipalpis Cook, New records. Djeboa ferruginea K. Djeboa unimaculata Cook, New records. Family Momoniidae K. Viets Subfamily Momonidinae Lundblad Genus Momonides Lundblad, A genus with a disjunct distribution, with two species known from the Oriental region, one from the Palaearctic region and one from the Afrotropical region. Momonides ghanaensis Smit, Figure 27 Figure Momonides ghanaensis Smit, female genital field.
Integument and lobed sclerites as described for male. Genital field with two genital plates, 88 long. Pre-genital sclerite 44 wide, post-genital sclerite 56 wide, more slender than pre-genital sclerite. Length of P1—5: 30, 50, 31, 44, Length of I-leg-4—6: 88, , Legs as in male. Thus far known only from the holotype male, and the female is described here for the first time. Atewa Hills, Ghana Smit References Bader C. Neue Fundorte von Wasssermilben aus dem Belgisch-Kongo.
Bader C. Wassermilben aus Zentralafrika. IN 8, Sci. Wassermilben aus Westafrika Acari, Prostigmata, Hydrachnellae. Cook D. Water mite genera and subgenera. New water mite species from tropical Africa Acari: Hydracarina. Zoologica, Stuttgart, Revisional study on water mites of the family Harpagopalpidae K. Viets, Acari: Hydrachnidia , with descriptions of new species from Cameroon and Madagascar.
Revisional studies on Thoracophoracarus K. Viets, Arachnida, Acari, Hydrachnidia, Arrenuridae. Zoosystema, Ecological studies on crater lakes in West Cameroon. Debundsha Lake. A new genus and species of water mite Acari: Hydrachnidia: Hygrobatidae from the prosobranch gastropod Lanistes libycus Morelet Streptoneura: Ampullariidae in Nigeria.
In: Smith, I. Indira Publishing House, West Bloomfield. Gledhill T. A new species of water mite Acari: Hydrachnidia: Hygrobatidae from the mantle cavity of the prosobranch Potadoma moerchi Reeve Streptoneura: Thiaridae in Nigeria. Zootaxa, Two new sympatric water-mites Acari: Hydrachnidia: Unionicolidae from the mutelid bivalve Aspatharia sinuata von Martens in Nigeria with some data on unionicoline - bivalve relationships.
A, Neue Serie, 3: Koenike F. Die Hydrachniden Ostafrikas. In: Stuhlmann F. Die Thierwelt Ost-Afrikas. Wirbellose Thiere, 6: Lundblad O. Afrikanische Hydracarinen. In: Explor. Park National Albert. Damas , Hydracarinen von den ostafrikanischen Gebirgen. The water mite family Mideopsidae Acari: Hydrachnidia : a contribution to the diversity in the Afrotropical region and taxonomic changes above species level. Torrenticolid water mites Acari: Hydrachnidia: Torrenticolidae from Ghana.
The invertebrate fauna of lake Tiberias. Qualitative aspects. Israel J. Smit H. New species of water mites from West Africa Acari, Hydrachnellae , including a checklist of species recorded from Cameroon. Storkia, 3: On the ground floor, a vast kitchen and dining area was created by knocking down an old conservatory and pushing the extension out by a further three metres, with floor-to-ceiling aubergine-coloured Crittall doors onto the garden. The six-month wait for planning permission for the extension turned out to be an unexpected bonus, giving the couple time to interrogate their plans.
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